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A little off topic, I always found it kind of weird though that Zetsu were supposed to be Hashirama clones, but the only successful Zetsus looked nothing like him.ya that would be unbelievable indeed but like you said, they were both connected, so it might be possible that they have a link.
Regarding your last paragraph, It's very interesting to see it from this perspective haha. They manipulated the hate inside Madara like madara tried to do with obito haha and instead of trying to bring peace like what they wanted, it shall bring destruction.
It could be that since Hashi cells r so close to the Shinjuu (both tree related) Hashi cells simply provided a proper medium in order to create Kaguya's servants.
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. So where does Spiral Zetsu get his access to the other 3 natures (wind, lightning, and fire)? Is it due to a more direct influence by the Mazou during his specific creation? Could this be showing that Spiral Zetsu has a deeper connection to the Shinju?