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Onoki and Muu are a prime example since Muu isn't even Onoki's brother; there is no bloodline connection between the two. It is extremely difficult to learn without the bloodline, but it can be done. Most likely with some very special methods requiring overwhelming skill.
So this shows why it is not a plothole for Prime Hiruzen to know all Konoha Jutsu, even all the KKGs. Afterall, he is the Professor.
Yes Mate. So with this logic anyone can learn Kimimaro's KKG, "Shikotsumyaku."
There is a difference from something that is directly passed down from your bloodline and something that can be learned (KKT). KKG cannot be taught, as they are passed down from your bloodline.